The introduction sequence introduces the 4 new Vault Hunters, lured on a train by Handsome Jack, as they are ambushed by Hyperion robots and the train is destroyed, leaving them for dead in the frozen wasteland. After waking, the surviving Vault Hunter is rescued by the last Claptrap unit and is taken to his home. The Guardian Angel contacts the new hunter and explains she is there to help and to keep the Claptrap unit safe and to head to Sanctuary, the capital of the resistance movement The Crimson Raiders. After acquiring a weapon the Vault Hunter is lead to Liar's Berg and meets with Sir Hammerlock. After helping Hammerlock, the Vault Hunter assists Claptrap in retrieving his ship from Captain Flynt, the local bandit leader. The Vault Hunter successfully defeats Flynt and with Claptrap in tow, heads for Sanctuary. After arriving, the Vault Hunter is tasked with proving his worth and successfully is inducted into the ranks of the Crimson Raiders. Entering Sanctuary, the Vault Hunter is then tasked with rescuing Roland, the leader of the Raiders. An Echo-communicator from the bounty hunter called the "Firehawk" demands the Vault Hunter to meet with him to acquire Roland. After fighting through many bandits, the Vault Hunter meets with Firehawk who is revealed to be Lilith, a previous Vault Hunter. Due to the introduction of large quantities of a substance known as Eridium, Lilith is now vastly more powerful siren and was tasked with keeping the bandit clans in check with Roland concentrated on the resistance. Lilith tells the Vault Hunter that the local bandits have kidnapped Roland and plan to sell him to Hyperion. Assaulting the bandit stronghold, the Vault Hunter meets Roland in his cell, but Hyperion's loaders attack the facility and retreat to extract Roland. Following the loaders, the Vault Hunter successfully rescues Roland. Accompanied by Lilith, the group heads back to Sanctuary to reassess the situation. Getting a information that the Vault Key will be transported by a local train, the group head to assault the train. Meeting up with former hunter Mordecai, and psychotic, orphaned 13-year-old Tiny Tina, the Vault Hunter is successfully derails the train. Upon attacking, the Vault Hunter encounters Hyperion's most powerful robot, Wilhelm. After defeating it, the Vault Hunter learns that the key was never on the train, but take Wilhelm's energy source as it could be useful. Roland tells the Vault Hunter to plug the energy source into Sanctuary's shield generator. However, upon plugging it in, Angel speaks with the player and apologizes before deactivating the shields. Handsome Jack tells the player he has been controlling Angel all along, even manipulating the events of the first game, to acquire the Vault Key. Upon rerouting power, Lilith uses her siren powers to activate the spaceship underneath Sanctuary, which prompts whole city floats to the sky. Lilith then teleports the city out of Handsome Jack's attack range. Upon reactivation of the Sanctuary's fast travel station, the player meets back with Roland. Meeting with Roland, Handsome Jack's plan is revealed. He wished to open a second Vault, this one containing a powerful warrior who is controlled by whomever opened the vault. Angel interrupts, and tells the group she can help, and tells them she has the Vault Key. It needs to be recharged every 200 years, but Jack has been using Angel and the Eridium to super charge the key. Angel tells them where the key is, but Hyperion has placed near-impenetrable defenses; a electrical wall scanner that can only be passed by a Hyperion robot, a massive bunker with enough ordnance to level a planet, and a final protocol that only Handsome Jack himself can disable. Roland figures that Claptrap can get past the scanner, and has the Vault Hunter meet up with Mordecai to get a software upgrade for him. Jack had kidnapped Bloodwing, Mordecai's best friend/pet/partner, and "slagged" her with a new element made from Eridum that Jack has been experimenting with. Jack kills Bloodwin.
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